Andrey Gebruk

Gebruk av

Acting Deputy Director for Scientific Work of Marine Ecology
Head of Laboratory
DSc (Biological Sciences)

Laboratory of Ocean Benthic Fauna
Marine Biology
Directorate of the IO RAS

36, Nakhimovskiy prospect, Moscow, 117997,
+7 (499)124-85-04

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Biologist, doctor of biological Sciences (2004), from 2002 – head of the Laboratory of benthic fauna of the ocean.

In 1982 he graduated from the Biological faculty of Moscow state University and in 1986 he defended his thesis on deep-sea holothurians. Since 1987 – he is the junior researcher of the laboratory of benthic fauna of the ocean, Institute of Oceanology.

Research interests are deep-water sea cucumbers and hydrothermal fauna.

In the first years of operation he participated in a series of expeditions to study shallow-water hydrothermal springs in the Western Pacific. In 1991 he made the first dive on the submersible "Mir" and since then regularly participated in studies hydrothermal fauna of the Mid-Atlantic ridge using deep-sea submersibles. In addition to DSS "Mir" he sank on devices "Pisis" and "Alvin".

In 1993-1995 he is a fellow of Alexander von Humboldt fund in the period of work at the Institute of hydrobiology and fisheries research of the University of Hamburg in the group of professor Hjalmar Thiel.

In 1995-1996 he is a fellow of the London Royal society, worked at the Oceanographic centre of Southampton (UK) in the group of professor Paul Tyler.

In 1997-1999 he worked in the Laboratory of the Marine biological Association in Plymouth (UK). Since 2002 he is the head of the Laboratory of benthic fauna of the ocean.

He is the author of new species and genera of deep-sea ba-legged holothurians. He has audited the elpidio families and floating holothurians of pelagothuriidae family. In 1990, according to the materials of a PhD thesis he published the monograph "the Deep-water holothurians of the elpidio family".

He is the author of a series of works on nutrition hydrothermal shrimps, one of the developers of the ideas about symbiotrophic diet of shrimps and versatility trophic structure of hydrothermal communities. Under his leadership, the community of the hydrothermal field Logachev (Mid–Atlantic ridge) and the mud volcano Haakon Mosby (Norwegian sea) was first described.

With his participation it was for the first time compiled and analyzed in a comparative perspective the information on the hydrothermal fauna of all known hydrothermal fields of the Atlantic ocean.

He has made a great contribution to the study of the biogeography of hydrothermal communities and taxa and vertical zonation. It is shown that the global distribution of hydrothermal fauna is subject to the same laws as the distribution of the ordinary, "dehydrothermal" fauna.

He maintains extensive international connections, he is a member of the scientific committees of the following international projects: "Euro Census of Marine Life", ChEss (Biogeography of Chemosynthetic Communities), АгсОD (Агсtic Ocean Diversity), МАR-EDC (Раtterns and processes of the ecosystems of the northern Mid-Atlantic). He is also included in the "InterRidhe Biology Working Group".

He is a member of the editorial boards of the journals "Marine Biology Research" and "invertebrate Zoology".

He is the author of about 100 publications, including 1 monograph. He was the editor in 2 books. He is the executive editor and co-author of the monograph "Biology of hydrothermal systems" (2002).

The most important publications

1983. Protelpidia new genus of the deep sea holothurians (Elasipodida, Elpidiidae). Zoologichesky Zhurnal (Zoological Journal) 62 (7): 1038 1043. (In Russian, English summary).

1983. Deep sea holothurians of the genus Scotoplanes (Elasipoda, Elpididae). Zool. Zhurnal 62 (9): 1359-1370. (In Russian, English summary).

1988. New taxa of the deep sea elpidiid holothurians. Zool.Zhurnal 67 (9): 914 922. (In Russian, English summary).

1988. Main types of sceletal elements of elasipodid holothurians. Zool. Zhurnal 67 (11): 1670 1679. (In Russian, English summary).

1989. Evolutionary trends of elasipodid holothurians. Trudy Instituta Okeanologii (123): 134 150. (In Russian).

1989. Revision of the family Pelagothuriidae (Holothurioidea, Elasipodida) with a review of swimming holothurians. Part 1. Revision of Pelagothuriidae. Zool. Zhurnal 68 (12): 57 66. (In Rusian, English summary).

1990. Revision of the family Pelagothuriidae with a review of swimming holothurians. Part 2. Review of swimming holothurians and their origin. Zool. Zhurnal 69 (3):60 68. (In Russian, English summary).

1990. About the record of pelagic larvae of elasipodid holothurians. Zool. Zhurnal 69 (8): 150 152. (In Russian, English summary).

1990. Deep sea holothurians of the family Elpidiidae. Moscow: Nauka. 160 p.

1991. Kuznetsov A.P., Gebruk A.V. Bacterial symbiotrophic feeding and organization of respiration feeding system in Axinopsida orbiculata orbiculata from Kraternaya Bay (Kuril Islands). Okeanologia 31 (1): 162 169.

1991. Zhukova N.V., Kharlamenko V.I., Gebruk A.V. Fatty acids of Axinopsida orbiculata - potential for detection of symbiosis with chemoauthotrophic bacteria. In: Shallow-water vents and ecosystem of Kraternaya Bight (Volcano Ushishir, Kuril Islands), Vol.1. Functional characteristics. Part II. DVO RAN, Vladivostok, pp. 63-78. (In Russian).

1992. The problem of "life-form" on the example of echinoderms. Uspechi sovremennoi biologii 112 (2): 176-185. (In Russian).

1992. Pimenov N.V., Savvichev A.S., Gebruk A.V., Moskalev L.I., Lein A.Yu., Ivanov M.V. Trophical specialization of Bresiliidae shrimps in the TAG site hydrothermal community. Doklady Academii Nauk 323 (3): 567-571. (In Russian).

1992. Gebruk, A.V., Pimenov N.V., Savvichev A.S. Bacterial "farm" on the mouthparts of shrimp. Priroda 6: 37-39. (In Russian).

1993. Vinogradova N.G., Belyaev G.M., Gebruk A.V. et al. Investigations of the Orkney Trench in the 43 cruise of the R.V. Dmitri Mendeleev. Geomorphology, sediments, benthic fauna. Trudy Instituta Okeanologii 127: 9-33. (In Russian).

1993. Vinogradova N.G., Gebruk A.V. et al. Abyssal benthic fauna of the Weddel Sea, the Scotia Sea and the Argentina basin on the results of the 43 cruise of the R.V. Dmitri Mendeleev. Trudy Instituta Okeanologii 127: 34-49. (In Russian).

1993. New records of elasipodid holothurians in the Atlantic sector of Antarctic and subantarctic. Trudy Instituta Okeanologii 127: 228-244. (In Russian).

1993. Gebruk A.V., Pimenov N.V., Savvichev A.S. Feeding specialization of Bresiliidae shrimps in the TAG site hydrothermal community. Mar. Ecol. Prog. Ser. 98: 247-253.

1993. Tarasov V.G.,... Gebruk A.V. Structural and functional pecularities of marine ecosystems in shallow-water zones of gasohydrothermal activity in Western Pacific Ocean. Izvestija Academii Nauk Ser. Biol. 6: 914-926. (In Russian).

1993. Gebruk A.V. et al. Role of bacterial component in feeding of deep-water bentic fauna in Froliha Bay, Lake Baikal. Izvestija Academii Nauk Ser. Biol. 6: 903-908. (In Russian).

Two main stages in the evolution of the deep-sea fauna of elasipodid holothurians. In: B. David, A. Guille, J.-P. Feral & M Roux (Eds.) Echinoderms through Time, Dijon : 507-514

1994. Revision of two deep-sea holothurian genera Psychrelpidia Herouard, 1923 and Rhipidothuria Herouard, 1901 (Elpidiidae). Mitt. hamb. zool. Mus. Inst., 91: 147-154.

1995. Locomotory organs in elasipodid holothurians: functional-morphological and evolutionary approaches. In: R. Emson, A. Smith & A. Campbell (eds) Echinoderm Research 1995. Balkema, Rotterdam, 95-102.

1996. Tyler, P.A., Rice, A.L., Young, C.M. & Gebruk A. A Walk on Deep Side: Animals in the Deep Sea. In: C.P.Summerhayes & S.A.Thorpe (Eds) Oceanography - An Illustrated Guide. Manson: London. pp. 195-211.

1997. New species of the deep-sea holothurian family Elpidiidae Th?el, 1879. Zoosystema, 19 (2-3), 211-217.

1997 Gebruk A.V., Tyler P.A. & Billett D.S.M. New records and review of pelagic juveniles of the deep-sea elasipodid holothurians. Ophelia, 46 (2), 153-164.

1997. Gebruk, A.V., Galkin, S.V., Vereshchaka, A.L., Moskalev, L.I. and Southward, A.J. Ecology and biogeography of the hydrothermal fauna of the Mid-Atlantic Ridge. 1997. Advances in Marine Biology, 32, 93-144.

1997. Lein, A.Y., Galkin, S.V., Gebruk, A.V., Pimenov, N.V., Miller, Y.V. , Moskalev L.I. and Ivanov, M.V. Role of bacterial chemosynhetsis in the nutrition of organisms in the Broken Spur hydrothermal community. Doklady Akademii Nauk, 357 (3), 410-413. [In Russian].

1998. Tarasov, V.G., Gebruk, A.V., Shulkin, V.M., Kamenev, G.M., Fadeev, V.I., Kosmynin, V.N., Malakhov, V.V., Starynin, D.A. and Obzhirov, A.I. Effect of shallow-water hydrothemal venting on the biota of Matupi Harbour (Rabaul Caldera, New Britain Island, Papua New Guinea). Continental Shelf Research 19(1), 79-116.

1998. Spicule changes during somatic growth in holothurians. In: Mooi, R. and Telford, M. (eds) Echinoderms: San Francisco, Balkema, Rotterdam, Brookfield, 456.

1998. Mironov, A.N., Gebruk, A.V. and Moskalev, L.I. Biogeographical patterns of the hydrothermal fauna: comparison with "non-vent biogeography". Cahiers de Biologie Marine, 39, 367-368.

1999. Богданов Ю.А., Сагалевич А.М., Вогт П.Р., Минерт Ю., Сандвор Э., Крейн К., Леин А.Ю., Егоров А.В., Пересыпкин В.И., Черкашев Г.А., Гебрук А.В., Гинсбург Г.Д., Войтов Д.В. Грязевой вулкан Хаакон Мосби в Норвежском море: результаты комплексных исследований с глубоководных обитаемых аппаратов. Океанология, 39, 412-419. [Bogdanov et al. 1999. Haakon Mosby Mud Volcano in the Norwegian sea: results of a complex study using deep diving submersibles. Okeanologiya, 39, 412-419].

1999. Bluhm H., and Gebruk A.V. Holothuroidea (Echinodermata) of the Peru Basin – Ecological and taxonomical remarks based on underwater images. P.S.Z.N. Marine Ecology, 20 (2), 167-195.

1999. Pimenov N.V., Savvichev A.S., Rusanov I., Lein, A.Y., Gebruk A.V. and Moskalev L.I. Microbial processes of carbon cycle as the base of the food chain of Haakon Mosby mud volcano benthic community. Geo-Marine Letters, 19 (1/2), 89-96.

1999. Luth U., Luth C., Gebruk A.V., Thiel H. Methane gas seeps along the oxic/anoxic gradient in the Black Sea: manifestations, biogenic sediment compounds, and preliminary results on benthic ecology. P.S.Z.N. Marine Ecology, 20(3-4), 221-249.

2000. Pond DW, Gebruk A, Southward EC, Southward AJ, Fallick AE, Bell MV, Sargent JR. Unusual fatty acid composition of storage lipids in the bresiliod shrimp, Rimicaris exoculata, couples the photic zone with MAR hydrothermal vent sites. Marine Ecology Progress Series, 198, 171-179.

2000. Smirnov A.V., Gebruk A.V., Galkin, S.V., Shank, T. New species of holothurian (Echinodermata, Holothuroidea) from hydrothermal vent habitats. Journal of the Marine Biological Association of the U.K., 80(2), 321-328.

2000.Gebruk, A.V., Chevaldonne, P., Shank, T., Lutz, R.A. and Vrijenhoek, R.C. Deep-sea hydrothermal vent communities of the Logatchev area (14o45’N, Mid-Atlantic Ridge): diverse biotopes and high biomass. Journal of the Marine Biological Association of the U.K., 80(3), 383-394.

2000. Gebruk, A.V., Southward, A.J., Southward, E.C., Kennedy, H. Food sources, behaviour and distribution of hydrothermal vent shrimps at the Mid-Atlantic Ridge. Journal of the Marine Biological Association of the U.K., 80(3), 487-499

2000. Roberts, D., Gebruk, A., Levin, V. and Manship, B.A.D. Feeding and digestive strategies in deposit-feeding holothurians. Oceanography and Marine Biology: Annual Review, 38, 257-310.

2001. Леин А.Ю., Пименов Н.В., Гебрук А.В., Москалев Л.И., Багиров Н.Э., Иванов М.В. Трофическая структура и состав гидротермального сообщества Рейнбоу (36 с.ш., Срединно-Атлантический хребет). Доклады РАН, 376 (1), 120-123. [Lein et al. 2001. Trophic structure of the Rainbow hydrothermal community (36 N, MAR). Doklady RAN, 376(1), 120-123].

2001. Southward, E.C., Gebruk, A., Kennedy, H., Southward, A., Chevaldonn? P. Differing energy sources for symbiotic bacteria of three bivalve molluscs at the Logatchev hydrothermal site (Mid-Atlantic Ridge). Journal of the Marine Biological Association of the U.K., 81(4), 655-661.

2002. Гебрук А.В., Галкин С.В. Гидротермальный биотоп и гидротермальная фауна: общие положения. Биология гидротермальных систем (Отв. ред. А.В. Гебрук), с. 13-24. Москва, КМК. [Gebruk A., Galkin S. 2002. Hydrothermal biotope and hydrothermal fauna: general remarks. In: A.V. Gebruk (ed) Biology of Hydrothermal Systems. Moscow, KMK, pp. 13-24].

2002. Верещака А.Л., Гебрук А.В. Гидротермальная фауна: состав, биология, адаптации. Креветки (Decapoda, Macrura, Natantia). Биология гидротермальных систем (Отв. ред. А.В. Гебрук), с. 185-197. Москва, КМК. [Vereshchaka A., Gebruk A. 2002. Hydrothermal fauna. Shrimps (Decapoda, Macrura, Natantia). In: A.V. Gebruk (ed) Biology of Hydrothermal Systems. Moscow, KMK, pp. 185-197].

2002. Гебрук А.В. Галки С.В., Леин А.Ю. Трофическая структура гидротермальных сообществ. В: Биология гидротермальных систем (Отв. ред. А.В. Гебрук), с. 351-362. Москва, КМК. [Gebruk A., Galkin S., Lein A. 2002. Trophic structure of hydrothermal vent communities. In: A.V. Gebruk (ed) Biology of Hydrothermal Systems. Moscow, KMK, pp. 351-362].

2002. Миронов А.Н., Гебрук А.В., Москалев Л.И. География гидротермальных сообществ и облигатных гидротермальных таксонов. Биология гидротермальных систем (Отв. ред. А.В. Гебрук), с. 410-455. Москва, КМК. [Mironov A., Gebruk A., Moskalev L. 2002. Biogeography of hydrothermal communities and obligate taxa. In: A.V. Gebruk (ed) Biology of Hydrothermal Systems. Moscow, KMK, pp. 410-455].

2003. Tunnicliffe, V., Baross, J., Gebruk, A.V., Giere, O., Koschinsky, A., Reysenbach, A.L., Shank, T., Summit, M. What are the interactions between biotic processes at vents and physical, chemical and geological conditions? In: Halbach P.E., Tunnicliffe V., Hein J.R. (eds) Energy and Mass Transfer in Marine Hydrothermal Systems. Dahlem Workshop Report 89. Berlin, Dahlem Univ. Press, pp. 251-270.

2003. A. V. Gebruk, H. Bluhm, T. Soltwedel, H. Thiel. Re-description of the enigmatic deep-sea holothurian Irpa abyssicola (Elpidiidae, Elasipodida) from the Norwegian-Greenland basin with notes on in situ observations. Sarsia, 88, 49-54.

2003. A.V. Gebruk, E.M. Krylova, A.Y. Lein, G.M. Vinogradov, E. Anderson, N.V. Pimenov, G.A. Cherkashev, K. Crane. Methane seep community of the H?kon Mosby mud volcano (the Norwegian Sea): Composition and trophic aspects. Sarsia, 88, 394-403.

2004. Гебрук А.В., Миронов А.Н. Проблемы изучения гидротермальной фауны в масштабе глобальных зон и периодов. В: Н.П. Лаверов (Отв. ред.). Актуальные проблемы океанологии. Москва, Наука, стр. 363-382. [Gebruk A., Mironov A. 2004. Hydrothermal vent fauna on a global scale in time and space. In: N. Laverov (ed) Actual problems of Oceanology. Moscow, Nauka, pp. 363-382].

2005. N.D. Holland, D.A. Clague, D.P. Gordon, A. Gebruk, D.L. Pawson, M.Vecchione. 2005. "Lophenteropneust" hypothesis refuted by collection and photos of new deep-sea hemichordates. Nature, 434, 17 March, 374-376.

Tarasov, V.G., Gebruk, A.V., Mironov, A.N., Moskalev, L.I. 2005. Deep-sea and shallow water hydrothermal vent communities: Two different phenomena? Chemical Geology, 224: 5-39.

Van Dover, C.L., Biscoito, M., Gebruk, A., Hashimoto, J., Tunnicliffe, V., Tyler, P. and Desbruy?res, D. 2006. Milestones in the discovery of hydrothermal-vent faunas. In:
Desbruy?res, D., Segonzac, M. and Bright, M. (Eds.) Handbook of Deep-Sea Hydrothermal Vent Fauna. Denisia, 18: 13–25.

Gebruk, A.V. 2006. Chiridota hydrothermica Smirnov et Gebruk, 2000. In: Desbruy?res, D., Segonzac, M. and Bright, M. (Eds.) Handbook of Deep-Sea Hydrothermal Vent Fauna. Denisia, 18: 480.

Галкин С.В., Гебрук А.В., Дехаирс Ф., Крылова Е.М., Леин А.Ю., Виноградов Г.М., Верещака А.Л. 2006. Трофическая структура североатлантических гидротерм: данные изотопного анализа. В кн.: Экосистемы атлантических гидротерм (Отв. ред. М.Е. Виноградов и А.Л. Верещака). Москва, Наука, стр. 95-117. [Galkin, S.V., Gebruk, A.V., Dehairs, F., et al. 2006. Trophic structure of North-Artlantic hydrothermal vents: isotopic data. In: M.E. Vinogradov and A.L. Vereshchaka (Eds). Ecosystems of Atlanrtic hydrothermal vents. Moscow: Nauka, pp.95-117].

Гебрук А.В., Миронов А.Н. 2006. Биогеография гидротерм Срединно-Атлантического хребта. В кн.: Экосистемы атлантических гидротерм (Отв. ред. М.Е. Виноградов и А.Л. Верещака). Москва, Наука, стр. 119-162. [Gebruk, A.V. and Mironov, A.N. 2006. Biogeography of Atlantic hydrothermal vents. In: M.E. Vinogradov and A.L. Vereshchaka (Eds). Ecosystems of Atlantic hydrothermal vents. Moscow: Nauka, pp.119-162].

Mironov A.N., Gebruk A.V. 2006. Biogeography of the Reykjanes Ridge, northern Atlantic. In: A.N. Mironov, A.V. Gebruk, A. J. Southward (Editors). Biogeography of the North Atlantic seamounts. Moscow: KMK Scientific Press Ltd., pp. 6-21.

2008. Bergstad, O.A., Falkenhaug, T., Astthorsson, O.S., Byrkjedal, I., Gebruk, A.V., Piatkowski, U., Priede, I.G., Santos, R.S., Vecchione, M., Lorance, P. and Gordon, J.D.M. Towards improved understanding of the diversity and abundance patterns of the mid-ocean ridge macro- and megafauna. Deep-Sea Research Part II, 55 (1-2): 1-5.

2008. Mortensen, P.B., Buhl-Mortensen, L., Gebruk, A.V. and Krylova, E.M. Occurrence of deep-water corals on the Mid-Atlantic Ridge based on MAR-ECO data. Deep-Sea Research Part II, 55 (1-2): 142-152.

2008. Gebruk, A.V. Holothurians (Holothuroidea, Echinodermata) of the northern Mid-Atlantic Ridge collected by the G.O. Sars MAR-ECO expedition with descriptions of four new species. Marine Biology Research, 4(1-2): 48-60.

2008. Budaeva, N.E., Mokievsky, V.O., Soltwedel, T. and Gebruk, A.V. 2008. Horizontal distribution patterns in Arctic macrobenthic deep-sea communities. Deep-Sea Research Part I, 55: 1167– 1178.

2009. Cross, I.A., Gebruk A., Rogacheva A., Billett D.S.M. 2009. Peniagone crozeti, a new species of elasipodid holothurian from abyssal depths off the Crozet Islands in the Southern Indian Ocean. Zootaxa 2096: 484-488.

Gebruk A.V., Budaeva N.E., King N.J. 2010. Bathyal benthic fauna of the Mid-Atlantic Ridge between the Azores and the Reykjanes Ridge. Journal of the Marine Biological Association of the United Kingdom, 2010, 90(1), 1–14.

Schander, C., Rapp, H.T., Bakken, T., Berge, J., Cochrane, S., Kongsrud, J.A., Oug, E., Byrkjedal, I., Cedhagen, T., Fosshagen, A., Gebruk, A., Larsen, K., Obst, M., Pleijel, F., St?hr, S., Todt, C., War?n, A., Hadler-Jacobsen, S., Keuning, R., Mikkelsen, N.T., Petersen, K.H., Torseth, I., Pedersen, R.B. 2010. The fauna of hydrothermal vents on the Mohn Ridge (North Atlantic). Marine Biology Research, 6 (2): 155-171.

Vecchione, M., Bergstad, O. A., Byrkjedal, I., Falkenhaug, R., Gebruk, A. V., God?, O. R., Gislason, A., Heino, M., H?ines, A. S., Menezes, G., Piatkowski, U., Priede, I. G., Skov, H., S?iland, H., Sutton, T., and de Lange Wenneck, T. 2010. Biodiversity Patterns and Processes on the mid-Atlantic Ridge. In: A. McIntyre (Ed.) Life in the World’s Oceans: Diversity, Distribution, and Abundance. Wiley Blackwell, Oxford, pp. 103–121.

Gradinger, R., Bluhm, B.A., Hopcroft, R.R., Gebruk, A.V., Kosobokova, K., Sirenko, B. And Weslawski, J.M. 2010. Marine Life in the Arctic. In: A.D. McIntyre (Ed.) Life in the World’s Oceans: Diversity, Distribution, and Abundance. Wiley Blackwell, Oxford, pp. 183–202.

Gebruk, A., Fabri, M.-C., Briand, P. and Desbruy?res, D. Community dynamics over a decadal scale at Logatchev, 14°45’N, Mid-Atlantic Ridge. 2010. Cahiers de Biologie Marine, 51: 383-388.

Fabri, M.-C., Bargain, A., Briand, P., Gebruk, A., Fouquet, Y., Morineaux, M. and Desbruy?res, D. The hydrothermal vent community of a new deep-sea field, Ashadze-1, 12°58’N on the Mid-Atlantic Ridge. 2011. Journal of Marine Biological Association of the United Kingdom, 91(1): 1-13.

Bluhm, B.A., Ambrose, W.G. Jr, Bergmann, M., Clough, L.M., Gebruk, A.V., Hasemann, C., Iken, K., Klages, M., MacDonald, I.R., Renaud, P.E., Schewe, I., Soltwedel, T. and Wlodarska-Kowalczuk, M. 2011. Diversity of the Arctic deep-sea benthos. Marine Biodiversity, 41: 87–107.

Bluhm, B.A., Gebruk, A.V., Gradinger, R., Hopcroft, R.R., Huettmann, F., Kosobokova, K.N., Sirenko, B.I. and Weslawski, J.M. 2011. Arctic Marine Biodiversity: An Update of Species Richness and Examples ?of Biodiversity Change. Oceanography 24(3): 232–248. 2.98

Krylova E. M., Gebruk A.V., Portnova D.A., Todt C., Haflidason H. 2011. New species of the genus Isorropodon (Bivalvia: Vesicomyidae: Pliocardiinae) from cold methane seeps at Nyegga (Norwegian Sea, Voring Plateau, Storrega slide), Journal of the Marine Biological Association of the United Kingdom, 91 (5), 1135-1144. 1.0

Jamieson, A.J., Gebruk, A., Fujii, T. and Solan, M. 2011. Functional effects of the hadal sea cucumber Elpidia atakama (Echinodermata: Holothuroidea, Elasipodida) reflect small-scale patterns of resource availability. Marine Biology, 158, (12), 2695-2703. (doi:10.1007/s00227-011-1767-7).

Gebruk A.V., Solis-Marin F.A., Billett D.S.M., Rogacheva A.V., Tyler P.A. 2012. Review of the genus Zygothuria Perrier, 1898 and the Atlantic group of species of the genus Mesothuria Ludwig, 1894 (Synallactidae: Holothuroidea) with description of the new species Mesothuria milleri sp. nov. Journal of Natural History, 46 (5–6): 265–348.

Karen J. Osborn, Linda A. Kuhnz, Imants G. Priede, Makoto Urata, Andrey V. Gebruk and Nicholas В. Holland. 2012. Diversification of acorn worms (Hemichordata, Enteropneusta) revealed in the deep sea. Proceedings of the Royal Society, B, Biological Sciences, 279, 1646–1654. doi: 10.1098/rspb.2011.1916.

Priede, I. G., Osborn, K. J., Gebruk, A. V., Jones, D., Shale, D., Rogacheva, A. and Holland, N. D. 2012. Observations on torquaratorid acorn worms (Hemichordata, Enteropneusta) from the North Atlantic with descriptions of a new genus and three new species. Invertebrate Biology, 131: 244–257.

Cross, I. A., Gebruk, A., Billett, D., Rogacheva, A. 2012. Rediscovery of the elpidiid holothurian Peniagone horrifer (Elasipodida, Holothuroidea: Echinodermata) in the southern Indian Ocean. Marine Biodiversity, 42, 241-245.

Rogacheva A.V., Gebruk A.V. and Alt C. 2013. Swimming deep-sea holothurians (Echinodermata: Holothuroidea) on the northern Mid-Atlantic Ridge. Zoosymposia, 7: 213-224.

Priede I.G., Bergstad O.A., Miller P.I., Vecchione M., Gebruk A., Falkenhaug T., Billett D.S.M., Craig J., Dale A.C., Shields M.A., Tilstone G.H., Sutton T.T., Gooday A.J., Inall M.E., Jones D.O.B., Martinez-Vicente V., Menezes G.M., Niedzielski T., Sigur?sson ?., Rothe N., Rogacheva A., Alt C.H.S., Brand T., Abell R., Brierley A.S., Cousins N.J., Crockard D., Hoelzel A.R., H?ines ?., Letessier T.B., Read J.F., Shimmield T., Cox M.J., Galbraith J.K., Gordon J.D.M., Horton T., Neat F., Lorance P. 2013. Does presence of a Mid Ocean Ridge enhance biomass and biodiversity? PLoS ONE 8(5): 1-10. doi:10.1371/journal.pone.0061550.

Gebruk A.V., Krylova E.M. 2013. Megafauna of the Charlie–Gibbs Fracture Zone (northern Mid-Atlantic Ridge) based on video observations. Journal of the Marine Biological Association of the United Kingdom, 93(5), 1143–1150. doi:10.1017/S0025315412001890.

Perez, J.A.A., E. dos Santos Alves, M.R. Clark, O. Aksel Bergstad, A. Gebruk, I. Azevedo Cardoso, and A. Rogacheva. 2012. Patterns of life on the southern Mid-Atlantic Ridge: Compiling what is known and addressing future research. Oceanography 25(4): 16–31. oceanog.2012.102.

Gebruk, A.V., Rogacheva, A.V., Pawson, D.L., Hamel, J.-F., Macisaac, K.G. & Mercier, A. (2013) Penilidia desbarresi sp. nov. (Echinodermata: Holothuroidea: Elasipodida) from the upper slope of Newfoundland and re-description of P. ludwigi (von Marenzeller, 1893). Marine Biology Research, 9, 1029–1036.

Rybakova (Goroslavskaya) E., Galkin S., Bergmann M., Soltwedel T., Gebruk A. 2013. Density and distribution of megafauna at the H?kon Mosby Mud Volcano (the Barents Sea) based on image analysis. Biogeosciences, 10: 3359–3374. doi:10.5194/bg-10-3359-2013.

Rogacheva A.V., Mironov A.N., Minin K.V., Gebruk A.V. 2013. Morphological evidence of depth-related speciation in deep-sea Arctic echinoderms. Invertebrate Zoology, 10 (1): 143–166.

Mironov A.N., Gebruk A.V., Dando P.R. Biogeography of the Arctic Ocean Thematic issue. Introduction. Invertebrate Zoology, 10 (1): 1-2.

Jones D.O.B., Alt C.H.S., Priede I.G., Reid W.D.K., Wigham B.D., Billett D.S.M., Gebruk A.V., Rogacheva A.V., Gooday A.J. 2013. Deep-sea surface-dwelling enteropneusts from the Mid-Atlantic Ridge: Their ecology, distribution and mode of life. Deep-Sea Research II 98: 374–387. (Thematic issue: ECOMAR: Ecosystems of the Mid-Atlantic Ridge at the Sub-Polar Front and Charlie-Gibbs Fracture Zone).

Osborn K.J., Gebruk A.V., Rogacheva A.V., Holland N.D. 2013. An Externally Brooding Acorn Worm (Hemichordata, Enteropneusta, Torquaratoridae) from the Russian Arctic. Biological Bulletin, 225: 113–123.

Rogacheva A., Gebruk A., Alt C.H.S. 2013. Holothuroidea of the Charlie Gibbs Fracture Zone area, northern Mid-Atlantic Ridge. Marine Biology Research, 9(5-6): 587-623.

Holland N.D., Osborn K.J., Gebruk A.V., Rogacheva A. 2013. Rediscovery and augmented description of the HMS ‘Challenger’ acorn worm (Hemichordata, Enteropneusta), Glandiceps abyssicola, in the equatorial Atlantic abyss. Journal of the Marine Biological Association of the United Kingdom, 93(8): 2197–2205. doi:10.1017/S0025315413000684.

2014. Gebruk A.V., Smirnov A.S., Rogacheva A.V. Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper 2000 m in seas bordering Europe. Holothuroidea. Invertebrate Zoology, 2014, 11(1): 156-180.

2016. Vedenin A., Budaeva N., Mokievsky V., Pantke C., Soltwedel T., Gebruk A. Spatial distribution patterns in macrobenthos along a latitudinal transect at the deep-sea observatory HAUSGARTEN. Deep-Sea Research Part I 114: 90–98.

2017. Molodtsova T., Galkin S., Kobyliansky S., Simakova U., Vedenin A., Dobretzova I., Gebruk A. First data on benthic and fish communities from the Mid-Atlantic Ridge, 16°40’-17°14N. Deep-Sea Research Part II, 137: 69–77.